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By: Ron, Santiago R. - Carrión, Julio - Caminer, Marcel - Sagredo, Yerka - Navarrete, María - Ortega, Jhael - Varela-Jaramillo, Andrea - Maldonado, Gabriela - Terán, ClaudiaType: ArticleIn: ZooKeysVolume: 993Series:Issue:Date: 2020Page Range: 121--155Publication Info: Pensoft Publishers 2020Subjects: Amazon Andes Brachycephaloidea Morphology phalanges Phylogeny Pristimantis amaguanae sp. nov Pristimantis nankints sp. nov Pristimantis romeroae sp. nov subarticular tubercles systematics taxonomy Terrarana