Biodiversity Data Journal 9: e62011 OO) doi: 10.3897/BDJ.9.e62011 open access Data Paper The Insect database in Dokdo, Korea: An updated version in 2020 Jihun Ryu*§, Young-Kun Kim!, Sang Jae Suh!, Kwang Shik Choi*S-1 $ School of Life Science, BK21 FOUR KNU Creative BioResearch Group, Kyungpook National University, Daegu, South Korea § Research Institute for Dok-do and Ulleung-do Island, Kyungpook National University, Daegu, South Korea | School of Applied Biosciences, Kyungpook National University, Daegu, South Korea 4] Research Institute for Phylogenomics and Evolution, Kyungpook National University, Daegu, South Korea Corresponding author: Kwang Shik Choi (ksc@knu.ac.kr) Academic editor: Paulo Borges Received: 14 Dec 2020 | Accepted: 20 Jan 2021 | Published: 26 Jan 2021 Citation: Ryu J, Kim Y-K, Suh SJ, Choi KS (2021) The Insect database in Dokdo, Korea: An updated version in 2020. Biodiversity Data Journal 9: e62011. https://doi.org/10.3897/BDJ.9.e62011 Abstract Background Dokdo, a group of islands near the East Coast of South Korea, comprises 89 small islands. These volcanic islands were created by an eruption that also led to the formation of the Ulleungdo Islands (located in the East Sea), which are approximately 87.525 km away from Dokdo. Dokdo is important for geopolitical reasons; however, because of certain barriers to investigation, such as weather and time constraints, knowledge of its insect fauna is limited compared to that of Ulleungdo. Until 2017, insect fauna on Dokdo included 10 orders, 74 families, 165 species and 23 undetermined species; subsequently, from 2018 to 2019, we discovered 23 previously unrecorded species and three undetermined species via an insect survey. New information As per our recent study, the database of insect species on Dokdo has been expanded to 10 orders, 81 families, 188 species and 23 undetermined species. This database has been © Ryu J et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. 2 Ryu J etal registered in the Global Biodiversity Information Facility (GBIF; www.GBIF.org) and is the first record for insect fauna on Dokdo. Keywords biodiversity, Dokdo, insect fauna, newly-recorded species, insect database Introduction Islands are known for their ecologically- and biologically-important ecosystems. Due to geographical isolation, the movement of organisms is limited (Franks 2009) and island-like areas have low potential for both species transfer and settlement. For these reasons, as well as their small land area, islands often have low biodiversity (Mauchamp 1997). Island- like areas are also vulnerable to external intrusions, which represent a major threat to indigenous species (Kil et al. 2006). Recent developments in traffic pathways have decreased the number of isolated island ecosystems, thus allowing researchers to investigate the interactions between evolutionary and ecological processes that are responsible for island biodiversity (Gillespie et al. 2008). The Intergovernmental Panel on Climate Change (IPCC) reports that climate change is causing sea levels and sea temperatures to rise and, if this trend continues, most coastal regions around the world will be at risk (Hong 2010, IPCC 2014). By 2100, these increases in sea level are estimated to reach at least 1 m, creating elevated flooding-related risks for large parts of the low-land island ecosystem, which could lead to significant habitat loss for many organisms worldwide (Bellard et al. 2013). The average annual sea level in Korean coastlines has been rising since 1989 and the average annual rate of sea level rise (5.67 mm/yr) around Ulleungdo has been particularly rapid (Korea Hydrographic and Oceanographic Agency 2020). The Dokdo Islands, which are located at 37°14'26.8" N and 131°52'10.4" E, belong to an administrative district that includes the Ulleungdo Islands as well. The address of the Korean administrative district is 1-96 Dokdo-ri, Ulleung-eup, Ulleung-gun, Gyeongsangbuk- do, Korea. Tourists cannot stay in Dokdo for more than 30 min, although the Korean Coast Guard (KCG) and some residents can visit for longer time periods. These Islands were formed during the Pliocene Epoch by an underwater volcanic eruption that occurred between 2.5 and 4.6 million years ago (Raman et al. 2016) and subsequently promoted the formation of the Ulleungdo volcanoes via tectonic plate movement (Lee et al. 2002). Dokdo is built on sea floor that is about 2,000 m deep; it comprises two main volcanic islands and 89 small islands (Sohn 1995, Ryu et al. 2012). The nearest land area to Dokdo is Jukbyeon, Uljin-gun, Gyeongsangbuk-do, which is 217.149 km away from Dokdo and 87.525 km from Ulleungdo (Hwang and Park 2007). Dokdo is entirely composed of volcanic rocks and its island ecosystem is relatively disconnected from the outside world; therefore, it is an important subject for island ecology The Insect database in Dokdo, Korea: An updated version in 2020 3 and biogeography (Cultural Heritage Administration 2009). In 1982, Dokdo was designated as Natural Monument No. 336 by the Korea Cultural Heritage Administration (KCHA) to be managed by the Dokdo Natural Reserve. Dokdo is very small, with an area of 187,554 m? and, because it has been protected and has inaccessible geographical features, there is extremely limited knowledge of its insect fauna. The Dokdo Islands are located at the bridge that connects Ulleungdo in Korea and the Oki Islands in Japan. Due to the fact that previous investigations of insect species on Dokdo played a vital role in characterising the biogeographic limits of these regions, the study of insect fauna on Dokdo is considered to be geographically important (Yasunaga and Duwal 2015). Since the initial assessment of insects on Dokdo by Jolivet in 1974, many researchers have conducted follow-up studies and, by 2017, 10 orders, 74 families and 165 species of insects have been identified (Jolivet 1974, Yoon 1978, Lee and Kwon 1981, Kwon et al. 1996, An 2000, Korean Ministry of Environment 2001, Ulleung Research Institute of Gyeongju University 2004, Park and Suh 2005, Kim and Yeom 2006, An 2008, Kim et al. 2009, Lee et al. 2009, Park et al. 2010, Park et al. 2011, Park et al. 2013, Daegu Regional Environmental Office 2012, Daegu Regional Environmental Office 2016, Choi et al. 2015, Park et al. 2017, Hwang et al. 2017). Here, we report 23 newly-identified and unrecorded species and three undetermined species on the Island and provide an updated database. Sampling methods Study extent: Between September 2017 and September 2018, we collected samples from Dokdo four times, once each at Anyongbok-gil, Ulleung-eup, Ulleung-gun and Gyeongsangbuk-do, Korea (131°52'03.2"E, 37°14'27.2'N), using sweeping, beating, brandishing, black light traps and pitfall traps. The survey was divided into East (Dongdo) and West (Seodo) using terrain isolation (Fig. 1). Figure 1. EESI Map of Dokdo: East (Dongdo) and West (Seodo) Islands. 4 Ryu J etal Sampling description: In Dongdo, we performed sample collection along the slope leading from the marina, through the KCG facility and then to the old marina. In Seodo, we performed sample collection along a very steep slope leading to the fishermen's dormitory. Seodo has less vegetation distribution compared to Dongdo. Collected specimen samples were stored in 70% ethanol in conical tubes. They were then transferred to the Animal Systematics and Taxonomy Laboratory and Pest Control Laboratory at Kyungpook National University, Korea. The samples were identified using the national species list of Korea and other references (National Institute of Biological Resources 2019, Chérot and Malipatil 2016, Yasunaga 2017, Gdllner-Scheiding 1976, Slater and Zheng 1984, Siwi and Doesburg 1984, Malenovsky 2013, ZHANG et al. 2013, Viraktamath 1979, Kim 2012, YOO et al. 2008, Yang et al. 2014, Kurahashi and Samerjai 2018, Suh and Kwon 2017a, Suh and Kwon 2017b, Bourquia et al. 2019, Vane-Wright and Hughes 2007, Ang and Meier 2010). Database update: We created a new checklist by adding 23 newly-confirmed insect species with reference to the previous reports and compiled it into a database. The data have been registered in the Global Biodiversity Information Facility (GBIF). Geographic coverage Description: The survey was divided into Dongdo and Seodo. Coordinates: 37-14 Latitude; 131-52 and 131-51 longitude. Taxonomic coverage Taxa included: Rank Scientific Name Common Name kingdom Animalia Animals phylum Arthropoda Arthropods class Insecta Insects order Blattodea family Ectobiidae order Coleoptera family Aphodiidae family Carabidae family Chrysomelidae family Coccinellidae The Insect database in Dokdo, Korea: An updated version in 2020 family Curculionidae family Elateridae family Endomychidae family Hydrophilidae family Latridiidae family Mordellidae family Nitidulidae family Oedemeridae family Scirtidae family Staphylinidae family Tenebrionidae order Dermaptera family Anisolabididae family Forficulidae order Diptera family Anthomyiidae family Calliphoridae family Ceratopogonidae family Chironomidae family Chloropidae family Coelopidae family Culicidae family Drosophilidae family Muscidae family Phoridae family Psychodidae family Rhiniidae family Sarcophagidae family Scathophagidae family Sepsidae family Syrphidae Tephritidae Tipulidae Hemiptera Alydidae Anthocoridae Aphididae Cicadellidae Cydnidae Delphacidae Lygaeidae Miridae Nabidae Pentatomidae Piesmatidae Rhopalidae Ryu J etal Rhyparochromidae Scutelleridae Tingidae Triozidae Hymenoptera Bethylidae Braconidae Chalcididae Eupelmidae Formicidae Ichneumonidae Pteromalidae Lepidoptera Crambidae Erebidae Geometridae The Insect database in Dokdo, Korea: An updated version in 2020 family Hesperiidae family Lycaenidae family Noctuidae family Nymphalidae family Papilionidae family Plutellidae family Pyralidae family Sphingidae family Tortricidae family Yponomeutidae order Neuroptera family Chrysopidae family Hemerobiidae family Sisyridae order Odonata family Aeshnidae family Coenagrionidae family Libellulidae order Orthoptera family Gryllacrididae family Gryllidae family Mogoplistidae Usage licence Usage licence: Creative Commons Public Domain Waiver (CC-Zero) Data resources Data package title: dokdo insect list, 1974-2019 Resource link: https://doi.org/10.15468/h684as Number of data sets: 1 Data set name: dokdo_insect_list_1974-2019 Ryu J etal Download URL: https:/Awww.gbif.org/dataset/eb47a0de-862f-44c0- be42-0e300abaaab4 Data format: CSV. Column label occurrencelD basisOfRecord eventDate institutionCode scientificName ScientificNameAuthorship collectionCode decimalLatitude decimalLongitude coordinateUncertaintyInMetres countryCode Identification Date Identified by stateProvince county locality catalogNumber vernacularName kingdom phylum class order family genus Column description Unique identifier of the occurrence State of the recorded specimen Date of the data registration Abbreviation of the institution having custody of the object Full scientific name The authorship information for the scientificName, formatted according to the conventions of the applicable nomenclaturalCode Abbreviation of specimen or database Geographic latitude of the collection site Geographic longitude of the collection site The horizontal distance (in metres) from the given decimalLatitude and decimalLongitude describing the smallest circle containing the whole of the Location Country code Date for identifying the specimen Identifier for the specimen Province in which the specimen was collected County in which the specimen was collected Locality in which the specimen was collected Specimen number for occurrence Common or vernacular name in Korea kingdom name phylum name class name order name family name genus name The Insect database in Dokdo, Korea: An updated version in 2020 9 Additional information Results and Discussion In this study, we first created an initial database, based on the results of surveys of insects on Dokdo from 1974 to 2017. This checklist confirmed the identification of 10 orders, 74 families, 165 species and 23 undetermined species of insects distributed on the Island. Through our recent sampling, we collected and identified 23 previously unrecorded insect species in Dokdo. Of these species, ten are in the Order Hemiptera (Leptocorisa chinensis, Batracomorphus diminuta, Macrosteles strifrons, Recilia coronifera, Geotomus convexus, Creontiades coloripes, Nesidiocons tenuis, Glaucias subpunctatus, Liorhyssus hyalinus and Horridipamera inconspicua); two are in the Order Coleoptera (Aphodius urostigma and Nacerdes melanura); ten are in the Order Diptera (Dasysyrphus bilineatus, Scaeva komabensis, Sepsis monostigma, Lucilia porphyrina, Stomorhina obsoleta, Sarcophaga brevicornis, Sarcophaga peregrina, Atherigona oryzae, Orchisia costata and Culicoides circumscriptus); and one is in the Order Lepidoptera (Vanessa indica) (Fig. 2). Figure 2. EESl Newly-recorded insect species on Dokdo Island. A. Creontiades coloripes; B.; Nesidiocoris tenuis; C. Liorhyssus hyalinus; D. Horridipamera inconspicua; E. Geotomus convexus; F. Glaucias subpunctatus; G. Leptocorisa chinensis; H. Recilia coronifera; |. Macrosteles striifrons; J. Batracomorphus diminuta; K. Nacerdes melanura; L. Dasysyrphus bilineatus; M. Scaeva komabensis; N. Sepsis monostigma; O. Lucilia porphyrina; P. Stomorhina obsoleta; Q. Sarcophaga brevicornis; R. Sarcophaga peregrina; S. Atherigona oryzae; T. Orchisia costata; U. Vanessa indica; V. Aphodius urostigma; W. Culicoides circumscriptus. Scale bars: F, G, L, M, R, U = 2.0 mm;A, C, D, K, O, P, Q= 1.0 mm; B, E, H, I, J, N, S, T, V, W= 0.5 mm. 10 Ryu J etal In addition to the 23 previously unrecorded species on Dokdo, three undetermined species were identified to the genus level: two Empoasca species in the Family Cicadellidae and one Sisyra species in the Family Sisyridae. Furthermore, five of the 23 previously unrecorded species and one of the three unidentified species belong to new families, adding a total of six newly-added families to the Dokdo Insect Database. Finally, we generated an updated database of insect fauna on Dokdo that contains 10 orders, 81 families, 188 species and 23 unidentified species (Table 1). Table 1. The updated database of insect species on Dokdo, based on data collected from 1974 to 2019. Order Family Scientific Name Newly recorded Undetermined species name Blattodea Ectobiidae Blattella nipponica Asahina, 1963 Coleoptera Aphodiidae Aphodius urostigma Harold, 1862 O Coleoptera Carabidae Anisodactylus signatus (Panzer, 1796) Coleoptera Carabidae Anisodactylus tricuspidatus A.Morawitz, 1863 Coleoptera Carabidae Dolichus halensis (Schaller, 1783) Coleoptera Carabidae Harpalus jureceki (Jedlicka, 1928) Coleoptera Carabidae Harpalus sinicus Hope, 1845 Coleoptera Carabidae Stenolophus difficilis (Hope, 1845) Coleoptera Chrysomelidae Callosobruchus chinensis (Linnaeus, 1758) Coleoptera Chrysomelidae Cassida nebulosa Linnaeus, 1758 Coleoptera Chrysomelidae Cassida piperata Hope, 1842 Coleoptera Chrysomelidae Longitarsus succineus (Foudras, 1859) Coleoptera Chrysomelidae Psylliodes punctifrons Baly, 1874 Coleoptera Chrysomelidae Thlaspida biramosa (Boheman, 1855) Coleoptera Coccinellidae Coccinella septempunctata Linnaeus, 1758 Coleoptera Coccinellidae Harmonia axyridis (Pallas, 1773) Coleoptera Coccinellidae Harmonia yedoensis (Takizawa, 1917) Coleoptera Coccinellidae Propylea japonica (Thunberg, 1781) Order Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera The Insect database in Dokdo, Korea: An updated version in 2020 Family Coccinellidae Coccinellidae Coccinellidae Curculionidae Curculionidae Curculionidae Curculionidae Curculionidae Curculionidae Curculionidae Curculionidae Curculionidae Elateridae Elateridae Elateridae Elateridae Endomychidae Endomychidae Hydrophilidae Latridiidae Latridiidae Mordellidae Mordellidae Nitidulidae Scientific Name Newly recorded Undetermined species name Scymnus babai Sasaji, 1971 Scymnus ferrugatus (Moll, 1785) Scymnus sp. Kugelann, 1794 O Auleutes sp. Dietz, 1896 O Cosmobaris scolopacea Germar, 1819 Ceutorhynchus albosuturalis (Roelofs, 1875) Dermestes tessellatocollis Motschulsky, 1860 Rhinoncus cribricollis Hustache, 1916 Rhinoncus jakovievi Faust, 1893 Scepticus insularis Roelofs, 1873 Scepticus uniformis Kono, 1930 Sitona lineatus (Linnaeus, 1758) Agrypnus miyamotoi (Nakane & Kishii, 1955) Melanotus castanipes (Paykull, 1800) Melanotus cete Candéze, 1860 Pectocera fortunei Candéze, 1873 Ancylopus melanocephalus (Olivier, 1808) Ancylopus pictus asiaticus (Strohecker, 1972) Hydrophilus acuminatus Motschulsky, 1854 Cortinicara gibbosa (Herbst, 1793) Stephostethus chinensis (Reitter, 1877) Mordella sp. Linnaeus, 1758 O Mordella tokejii Nomura, 1958 Omosita colon (Linnaeus, 1758) 11 12 Order Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Coleoptera Dermaptera Dermaptera Dermaptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Family Nitidulidae Oedemeridae Scirtidae Staphylinidae Staphylinidae Staphylinidae Staphylinidae Staphylinidae Staphylinidae Tenebrionidae Tenebrionidae Anisolabididae Anisolabididae Forficulidae Anthomyiidae Anthomyiidae Anthomyiidae Anthomyiidae Calliphoridae Calliphoridae Calliphoridae Calliphoridae Calliphoridae Ceratopogonidae Chironomidae Ryu J etal Scientific Name Newly recorded species Omosita japonica Reitter, 1874 Nacerdes melanura (Linnaeus, 1758) O Cyphon sp. Paykull, 1799 Aleochara fucicola Sharp, 1874 Atheta sp. Thomson, 1858 Atheta tokiokai (K.Sawada, 1971) Cafius histrio (Sharp, 1874) Neobisnius sp. Ganglbauer, 1895 Paederus fuscipes Curtis, 1826 Gonocephalum coenosum Kaszab, 1952 Gonocephalum coriaceum Motschulsky, 1857 Anisolabis maritima (Bonelli, 1832) Euborellia annulipes (H.F.Lucas, 1847) Forficula scudderi de Bormans, 1880 Delia platura (Meigen, 1826) Fucellia apicalis Kertesz, 1908 Fucellia boninensis Snyder, 1965 Pegomya cunicularia (Rondani, 1866) Calliphora nigribarbis Smellen van Vollenhoven, 1863 Hemipyrellia ligurriens (Wiedemann, 1830) Lucilia illustris (Meigen, 1826) Lucilia porphyrina (Walker, 1856) O Lucilia sericata (Meigen, 1826) Culicoides circumscriptus Kieffer, O 1918 Polypedilum sp. Kieffer, 1912 Undetermined name Order Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera The Insect database in Dokdo, Korea: An updated version in 2020 Family Chloropidae Coelopidae Culicidae Culicidae Drosophilidae Muscidae Muscidae Muscidae Muscidae Phoridae Psychodidae Psychodidae Rhiniidae Sarcophagidae Sarcophagidae Sarcophagidae Scathophagidae Sepsidae Syrphidae Syrphidae Syrphidae Syrphidae Syrphidae Syrphidae Syrphidae Scientific Name Newly recorded Undetermined species name Thaumatomyia notata (Meigen, 1830) Coelopa frigida (Fabricius, 1805) Culex orientalis Edwards, 1921 Ochlerotatus togoi (Theobald, 1907) Drosophila sp. Fallén, 1823 O Atherigona oryzae Malloch, 1925 O Musca bezzii Patton & Cragg, 1913 Musca hervei Villeneuve, 1922 Orchisia costata (Meigen, 1826) O Megaselia spiracularis Schmitz, 1938 Psychoda alternata Say, 1824 Tinearia alternata (Say, 1824) Stomorhina obsoleta Wiedemann, O 1830 Sarcophaga brevicornis Ho, 1934 O Sarcophaga melanura Meigen, 1826 Sarcophaga peregrina (Robineau- O Desvoidy, 1830) Scathophaga stercoraria (Linnaeus, 1758) Sepsis monostigma Thomson, 1869 O Allobaccha apicalis (Loew, 1858) Allograpta javana (Wiedemann, 1824) Betasyrphus serarius (Wiedemann, 1830) Dasysyrphus bilineatus (Matsumura, O 1917) Episyrphus balteatus (De Geer, 1776) Eristalis cerealis Fabricius, 1805 Eristalis tenax (Linnaeus, 1758) 13 14 Order Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Diptera Hemiptera Hemiptera Hemiptera Hemiptera Hemiptera Hemiptera Hemiptera Hemiptera Hemiptera Hemiptera Hemiptera Hemiptera Hemiptera Family Syrphidae Syrphidae Syrphidae Syrphidae Syrphidae Syrphidae Tephritidae Tephritidae Tephritidae Tephritidae Tipulidae Alydidae Anthocoridae Anthocoridae Aphididae Aphididae Cicadellidae Cicadellidae Cicadellidae Cicadellidae Cicadellidae Cicadellidae Cicadellidae Cicadellidae Ryu J etal Scientific Name Newly recorded species Metasyrphus corollae (Fabricius, 1794) Metasyrphus nitens (Zetterstedt, 1843) Scaeva komabensis (Matsumura, O 1918) Melanostoma mellinum (Linnaeus, 1758) Sphaerophoria menthastri (Linnaeus, 1758) Xanthandrus comtus (Harris, 1780) Campiglossa sada (Dirlbek & Dirlbekova, 1974) Campiglossa sp. Rondani, 1870 Ensina sonchi (Linnaeus, 1767) Trupanea convergens (Hering, 1936) Tipula sp. Linnaeus, 1758 Leptocorisa chinensis Dallas, 1852 O Orius sauteri (Poppius, 1909) Orius sp. Distant, 1904 Aphis nerii Fonscolombe, 1841 Aphis rumicis Linnaeus, 1758 Balclutha rubrinervis Matsumura, 1902 Batracomorphus diminuta O Matsumura, 1912 Empoasca sp.1 Walsh, 1862 Empoasca sp.2 Walsh, 1862 Hishimonus sellatus Uhler, 1896 Laburrus impictifrons Boheman, 1852 Macrosteles striifrons Anufriev, 1968 O Maiestas oryzae Matsumura, 1902 Undetermined name The Insect database in Dokdo, Korea: An updated version in 2020 Order Family Scientific Name Newly recorded Undetermined species name Hemiptera Cicadellidae Psammotettix striatus Linnaeus, 1758 Hemiptera Cicadellidae Recilia coronifera (Marshall, 1866) O Hemiptera Cydnidae Geotomus pygmaeus (Dallas, 1851) Hemiptera Cydnidae Geotomus convexus Hsiao, 1977 O Hemiptera Delphacidae Laodelphax striatellus (Fallén, 1826) Hemiptera Delphacidae Sogatella furcifera (Horvath, 1899) Hemiptera Delphacidae Sogatella kolophon (Kirkaldy, 1907) Hemiptera Delphacidae Unkanodes sapporonus (Matsumura, 1935) Hemiptera Lygaeidae Nysius plebeius Distant & W.L., 1883 Hemiptera Miridae Campylomma lividicornis Reuter, 1912 Hemiptera Miridae Campylomma sp. Reuter, 1878 O Hemiptera Miridae Creontiades coloripes Hsiao & Meng, O 1963 Hemiptera Miridae Nesidiocoris tenuis (Reuter, 1895) O Hemiptera Miridae Orthotylus flavosparsus (C.Sahlberg, 1841) Hemiptera Miridae Trigonotylus caelestialium (Kirkaldy, 1902) Hemiptera Nabidae Prostemma hilgendorfii Stein, 1878 Hemiptera Pentatomidae Glaucias subpunctatus (Walker, O 1867) Hemiptera Pentatomidae Nezara antennata Scott, 1874 Hemiptera Piesmatidae Piesma capitatum (Wolff, 1804) Hemiptera Piesmatidae Piesma maculatum (Laporte, 1833) Hemiptera Rhopalidae Liorhyssus hyalinus (Fabricius, 1794) O Hemiptera Rhyparochromidae_ Horridipamera inconspicua Dallas, O 1852 Hemiptera Rhyparochromidae Paradieuches dissimilis (Distant & W.L., 1883) 16 Order Hemiptera Hemiptera Hemiptera Hemiptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Hymenoptera Family Rhyparochromidae Scutelleridae Tingidae Triozidae Bethylidae Braconidae Braconidae Braconidae Braconidae Braconidae Chalcididae Chalcididae Eupelmidae Eupelmidae Formicidae Formicidae Formicidae Formicidae Formicidae Formicidae Formicidae Formicidae Formicidae Formicidae Formicidae Ryu J etal Scientific Name Newly recorded species Stigmatonotum rufipes (V.Motschulsky, 1866) Cantao ocellatus (Thunberg, 1784) Cantacader lethierryi Scott, 1874 Trioza chenopodi Reuter, 1876 Acrepyris minutus Yasumatsu, 1955 Apanteles sp. Férster, 1862 Cotesia sp.1 Cameron, 1891 Cotesia sp.2 Cameron, 1891 Deuterixys sp. Mason, 1981 Lysiphlebus sp. Férster, 1862 Brachymeria femorata (Panzer, 1801) Brachymeria minuta (Linnaeus, 1767) Eupelmus australiensis (Girault, 1913) Eupelmus sp. Dalman, 1820 Hypoponera nippona (Santschi, 1937) Lasius meridionalis (Bondroit, 1920) Monomorium floricola (Jerdon, 1851) Monomorium intrudens Smith, 1874 Myrmecina graminicola nipponica Wheeler, 1906 Pachyconadyla chinensis (Emery, 1895) Pheidole fervida Smith, 1874 Ponera japonica Wheeler, 1906 Pristomyrmex pungens Mayr, 1866 Solenopsis japonica Wheeler, 1928 Strumigenys lewisi Cameron, 1886 Undetermined name Order Hymenoptera Hymenoptera Hymenoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera Lepidoptera The Insect database in Dokdo, Korea: An updated version in 2020 Family Formicidae Ichneumonidae Pteromalidae Crambidae Crambidae Crambidae Crambidae Crambidae Erebidae Geometridae Geometridae Hesperiidae Lycaenidae Lycaenidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Noctuidae Nymphalidae Nymphalidae Papilionidae Plutellidae Scientific Name Newly recorded Undetermined species name Tetramorium caespitum (Linnaeus, 1758) Homotropus sp. Férster, 1869 O Halticoptera circulus (Walker, 1833) Cnaphalocrocis medinalis Guenée, 1854 Diaphania indica (Saunders, 1851) Maruca vitrata (Fabricius, 1787) Palpita nigropunctalis Bremer, 1864 Spoladea recurvalis (Fabricius, 1775) Catocala dula Bremer, 1861 Odontopera arida Butler, 1878 Scopula ignobilis Warren, 1901 Parnara guttatus Bremer & Grey, 1853 Arhopala bazalus (Hewitson, 1862) Pseudozizeeria maha (Kollar, 1844) Agrotis ipsilon (Hufnagel, 1766) Arcte coerula Guenée, 1852 Callopistria argyrosticta Butler, 1881 Cosmia achatina Butler, 1879 Daddala lucilla Butler, 1881 Diarsia canescens Butler, 1878 Thyas juno (Dalman, 1823) Macdunnoughia confusa (Stephens, 1850) Mythimna separata (Walker, 1865) Cynthia cardui Linnaeus, 1758 Vanessa indica Herbst, 1794 O Papilio xuthus Linnaeus, 1767 Plutella xylostella (Linnaeus, 1758) 1904) 18 Ryu J etal Order Family Scientific Name Newly recorded Undetermined species name Lepidoptera Pyralidae Oncocera semirubella (Scopoli, 1763) Lepidoptera Sphingidae Macroglossum stellatarum (Linnaeus, 1758) Lepidoptera Tortricidae Adoxophyes orana (Fischer v.Roslerstamm, 1834) Lepidoptera Tortricidae Archips oporana (Linnaeus, 1758) Lepidoptera _ Tortricidae Cochylidia contumescens Meyrick, 1931 Lepidoptera _ Tortricidae Cochylidia richteriana (Fischer v.Roslerstamm, 1837) Lepidoptera _ Tortricidae Tortrix sinapina Butler, 1879 Lepidoptera Yponomeutidae Yponomeuta meguronis Matsumura, 1931 Neuroptera Chrysopidae Chrysopa pallens (Rambur, 1838) Neuroptera Hemerobiidae Hemerobius humulinus Linnaeus, 1758 Neuroptera _— Sisyridae Sisyra sp. Burmeister, 1839 O Odonata Aeshnidae Anax parthenope Selys, 1839 Odonata Coenagrionidae Ischnura asiatica Brauer, 1865 Odonata Libellulidae Pantala flavescens Fabricius, 1798 Odonata Libellulidae Rhyothemis fuliginosa Selys, 1883 Odonata Libellulidae Sympetrum darwinianum Selys, 1883 Orthoptera Gryllacrididae Nippancistroger sp. Griffini, 1913 O Orthoptera Gryllidae Teleogryllus emma (Ohmachi & |.Matsuura, 1951) Orthoptera Gryllidae Velarifictorus aspersus (F.Walker, 1869) Orthoptera Mogoplistidae Ornebius kanetataki (S.Matsumura, Amongst the 23 previously unrecorded species on Dokdo, B. diminuta was first discovered in Korea and the remainder of this species has been recorded in the Korean Peninsula. In addition, C. circumscriptus, a blood-sucking insect known for being extremely annoying to The Insect database in Dokdo, Korea: An updated version in 2020 19 the KCG and residents of Dokdo, has been identified for the first time in Dokdo, using a black light trap with dry ice. 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