JHR 84: I-9 (2021) re JOURNAL OF | *reerrtievedopenacoss ural dot: 103897/84.7212 (f-) Hymenoptera https://jhr.pensoft.net The Imernational Society of Hymenoperiss. RESEARCH Editorial Maxim Yu. Proshchalykin', Vladimir E. Gokhman? I Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok 690022, Russia 2. Botanical Garden, Moscow State University, Moscow 119234, Russia Corresponding author: Maxim Yu. Proshchalykin (proshchalikin@biosoil.ru) Academic editor: Michael Ohl | Received 27 July 2021 | Accepted 30 July 2021 | Published 24 August 2021 http://zoobank. org/08D07940-A5BC-4188-A434-06B2EEBIE536 Citation: Proshchalykin MYu, Gokhman VE (2021) Editorial. In: Proshchalykin MYu, Gokhman VE (Eds) Hymenoptera studies through space and time: A collection of papers dedicated to the 75" anniversary of Arkady S. Lelej. Journal of Hymenoptera Research 84: 1-9. https://doi.org/10.3897/jhr.84.72212 On August 25, 2021, Arkady S. Lelej, an outstanding Russian entomologist and a very fine person, turns 75. In addition to their warmest regards to him, Arkady’s colleagues and friends prepared a Special Issue of the Journal of Hymenoptera Research on this spe- cial occasion. 53 authors from Belgium, China, Czech Republic, Estonia, Finland, Ger- many, Italy, Japan, the Netherlands, Trinidad and Tobago, Ukraine, United Kingdom, USA, and Russia (Feodosiya, Kaliningrad, Moscow, Nizhny Novgorod, Petrozavodsk, Saratov, St. Petersburg, and Vladivostok) took part in preparing this Issue. They have jointly presented 24 papers dedicated to various aspects of Hymenoptera studies, includ- ing descriptions of five insect genera, 28 species and one subspecies as new to science. Among them, 18 new taxa (three genera and 15 species) are named in Arkady’s honor. Most papers in this Special Issue are dedicated to taxonomy and faunistics of vari- ous hymenopteran taxa. These include members of all traditional major subdivisions of the order, i.e., Symphyta, Parasitica and Aculeata. Among the extant taxa, there are members of the West Palaearctic Euura bergmanni and oligospila species groups (Ten- thredinidae) (Prous et al. 2021), Ichneumonidae from the Eastern Palearctic and Ori- ental regions (Humala 2021; Kasparyan and Khalaim 2021), Afrotropical members of the braconid genus Sculptolobus Yang et al., 2008 (Samartsev and van Achterberg 2021), chalcid wasps of the genus Toxeuma Walker, 1833 (Pteromalidae) of the Rus- sian fauna (Tselikh and Dale-Skey 2021), flat wasps of the genus Sierola Cameron, 1881 (Bethylidae) from China (Wang et al. 2021), and a new species of the genus Odontosphex Arnold, 1951 (Crabronidae) from Central Asia (Mokrousov and Prosh- Copyright Maxim Yu. Proshchalykin, Vladimir E. Gokhman. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. 2 M.Yu. Proshchalykin & V.E. Gokhman / Journal of Hymenoptera Research 84: 1-9 (2021) Ste ates ee at oe, a hee ¢ . } se wu : 4 ek / ae * ie! 1 7 i! { >. oe by as | j cP Sues 3) Fi Figure |. 9° International Congress of Hymenopterists. Ehime University, Matsuyama, Japan, 23-27 July 2018. Arkady S. Lelej presents his talk at the Congress. 26 July 2018. chalykin 2021), a new species of the genus Anteon Jurine, 1807 (Dryinidae) from Turkey (Speranza et al. 2021) as well as cuckoo wasps (Chrysididae) from Pakistan and India (Rosa et al. 2021), two bee genera from the Caucasus, Mongolia, Middle East and North Africa (Astafurova and Proshchalykin 2021; Proshchalykin and Dathe 2021; Bogusch 2021) and new genera and species of spider wasps (Pompilidae) and velvet ants (Mutillidae) from the Oriental Region (Shimizu et al. 2021; Loktionov 2021; Okayasu 2021). In addition, a few extinct groups were described, mostly from Baltic, Rovno and Sakhalinian ambers. These taxa include braconid parasitoids of ants from the tribe Neoneurini (Belokobylskij et al. 2021) as well as new genera and species of Encyrtidae (Simutnik et al. 2021) and Formicidae (Radchenko and Proshchalykin 2021). Moreo- ver, a new genus and species of the extinct family Peleserphidae (Proctotrupoidea) are described by Rasnitsyn (2021). In a single morphological study, the ultrastructure of unusual placoid sensilla of miniature Megaphragma Timberlake, 1924 wasps (Trichogrammatidae) was examined with various methods of electron microscopy (Diakova and Polilov 2021). Editorial 3 NIVERSITY 8) > BALENCIAGA PRIS i ee pete Wileceba ae Figure 2. 9" International Congress of Hymenopterists. Participants from Russia (left to right): Andrey I. Khalaim, Arkady S. Lelej, Ekaterina V. Tselikh, Vladimir E. Gokhman. 23 July 2018. Several papers are dedicated to the bionomics of certain groups of Hymenoptera, including details of parasitism by two Eupelmus Dalman, 1820 species (Eupelmidae) on Aulacidea hieracii (Bouché, 1834) (Cynipidae) (Gokhman and Nikelshparg 2021), additional nesting data for Leptochilus regulus (de Saussure, 1855) (Vespidae) (Fateryga and Fateryga 2021) as well as defensive adaptations of Mutillidae (Schmidt et al. 2021) and caste and wing loading in Dolichovespula maculata (Linnaeus, 1763) (Vespidae) (Starr et al. 2021). We celebrated Arkady’s 70" anniversary just five years ago. On that occasion, a Special Issue of the Euroasian Entomological Journal was published (http://www. eco.nsc.ru/eej/supplement_2016_15_1.html ). In this Issue, Arkady’s biography, together with a full list of his published papers and taxa described by him, is given. Nevertheless, Arkady did not decrease his productivity since that time. He contin- ues to work intensively, coupling research with organizational activities. Arkady continues to lead the Laboratory of Entomology of the Federal Scientific Center of 4 M.Yu. Proshchalykin & V.E. Gokhman / Journal of Hymenoptera Research 84: 1-9 (2021) the East Asia Terrestrial Biodiversity in Vladivostok. He is the Head of the Publish- ing Commission, and a member of both the Scientific Council and Dissertation Panel of the Center. During the last five years, Arkady led three projects funded by the Russian Foundation for Basic Research (RFBR), including IV Eurasian Sympo- sium on Hymenoptera (September 9-15, 2019, Vladivostok). Starting from 2017, he published more than 50 papers dedicated to various hymenopteran taxa as well as to members of other insect orders, in addition to about 350 papers published during the previous years. List of selected papers authored by Arkady Lelej (2017 — June 2021) 2017 Brothers DJ, Lelej AS (2017) Phylogeny and higher classification of Mutillidae (Hy- menoptera) based on morphological reanalyses. Journal of Hymenoptera Research 60: 1-97; Zhou H-T, Lelej AS, Xu Z-F (2017) Discovery of the subfamily Rhopalomutillinae (Hymenoptera: Mutillidae) in China and description of Pherotilla brothersi sp. nov. Zootaxa 4273(4): 587-594. Lelej AS, Williams KA, Loktionov VM, Pang H, Xu Z (2017) Review of the genus Zeugomutilla Chen, 1957 (Hymenoptera, Mutillidae, Mutillini), with description of two new species. Zootaxa 4247(1): 1-15. Lelej AS, Proshchalykin MYu, Loktionov VM (Eds) (2017) Annotated catalog of the Hymenoptera of Russia. Volume I. Symphyta and Aculeata. Proceedings of the Zoological Institute RAS, Supplement 6: 5-475. Loktionov VM, Lelej AS, Xu Z-F (2017) Review of the Priocnemis hyalinata species- group (Hymenoptera: Pompilidae), with description of a new species from China. Zootaxa 4254: 551-562. Loktionov VM, Lelej AS, Xu Z-F (2017) Review of the genus Eopompilus Gussako- vskij, 1932 (Hymenoptera: Pompilidae) with the description of new species from China. Zootaxa 4277: 413-426. Loktionov VM, Lelej AS, Xu Z-F (2017) Discovery of the genus Nipponodipogon Ishi- kawa in the Oriental region, with description of two new species from China (Hy- menoptera, Pompilidae). ZooKeys 692: 103-127. Sakhvon VV, Lelej AS (2017) Review of the genus Habropogon Loew, 1847 (Dip- tera: Asilidae) from Russia, with description of new species from Siberia. Zootaxa 4311(3): 417-425. Lelej AS, Mokrousov MV (2017) The types of Scoliidae (Hymenoptera), described by Eduard Eversmann, with some taxonomic notes and checklist of Russian Scoliidae. Far Eastern Entomologist 340: 1-17. Rosa P, Lelej AS, Proshchalykin MYu, Loktionov VM, Mokrousov MV (2017) New records of Chrysididae (Hymenoptera) from Russia with description of five new species. Far Eastern Entomologist 345: 1-33. Editorial 5 2018 Barthélémy C, Lelej AS, Loktionov VM (2018) A contribution to the Mutillidae (Hy- menoptera) of Hong Kong. Zootaxa 4514(2): 230-242. Zhou H-T, Lelej AS, Williams KA, Liu J-X (2018) Revision of the Oriental genus Zavatilla Vsuneki (Hymenoptera, Mutillidae, Trogaspidiini), with descriptions of two new species. Zootaxa 4418(2): 101-120. Okayasu J, Williams KA, Lelej AS (2018) A remarkable new species of Sinotilla Lele} (Hymenoptera: Mutillidae: Smicromyrmini) from Taiwan and an overview of color diversity in East Asian mutillid females. Zootaxa 4446(3): 301-324. Mokrousov MV, Lelej AS, Fadeev KI (2018) New data on the Palaearctic wasps of the genus Sierolomorpha Ashmead, 1903 (Hymenoptera, Sierolomorphidae) with description of S. trjapitzini sp. n. from Primorskii Territory, Russia. Entomological Review 98(6): 737-742. Loktionov VM, Lelej AS (2018) Discovery of the spider wasp genus Sphictostethus Kohl, 1884 (Hymenoptera: Pompilidae: Pepsinae) in New Guinea, with descrip- tion of two new species. Zootaxa 4429: 295-302. Loktionov VM, Lelej AS (2018) To the knowledge of genus Nipponodipogon Ishikawa, 1965 (Hymenoptera: Pompilidae, Pepsinae) from Laos. Far Eastern Entomologist SOP S17., Loktionov VM, Lelej AS, Liu J-X (2018) Contribution to the knowledge on the spider wasps genus Morochares Banks, 1934 (Hymenoptera: Pompilidae) from China. Zootaxa 4462: 497-510. Mokrousov MV, Lelej AS, Gorbatovsky VV (2018) The types of described by Ed- uard Eversmann Tiphiidae and Sierolomorphidae (Hymenoptera) with taxonomic notes. Far Eastern Entomologist 349: 11-16. Zhou H-T, Lelej AS, Liu J-X (2018) New records of velvet ants (Hymenoptera: Mu- tillidae) from China with taxonomic notes. Far Eastern Entomologist 354: 23-28. Shimizu A, Lelej AS, Loktionov VM, Nishimoto Y, Endo T (2018) Revision of the subgenus Stigmatodipogon Ishikawa of the genus Dipogon Fox (Hymenoptera: Pompilidae: Pepsinae). Zootaxa 4514: 1-22. Wahis R, Lelej AS, Loktionov VM (2018) Contribution to the knowledge of the genus Eopompilus Gussakovskij, 1932 (Hymenoptera, Pompilidae). Far Eastern Ento- mologist 361: 1-11. Sakhvon VV, Lelej AS (2018) Review of the genus Heteropogon Loew, 1847 (Dip- tera: Asilidae) from Russia, and Central Asia, with description of two new species. Zootaxa 4486(4): 435-450. 2019 Achterberg C, van, Sundukov YuN, Lelej AS, Belokobylskij SA (2019) To the knowl- edge of Gasteruptiidae (Hymenoptera) from Russian Far East. Far Eastern Ento- mologist 381: 1-8. 6 M.Yu. Proshchalykin & V.E. Gokhman / Journal of Hymenoptera Research 84: 1-9 (2021) Brothers DJ, Lelej AS, Williams KA (2019) Genus-group names of Mutillidae (Hyme- noptera): corrections and updates since 2008. Zootaxa 4651(3): 578-588. Kochetkov DN, Olmi M, Lelej AS (2019) Contribution to the knowledge of dryinid wasps genus Anteon Jurine, 1807 (Hymenoptera: Dryinidae) in the Russian Far East. Far Eastern Entomologist 323: 11-17. Williams KA, Lelej AS, Thaochan N (2019) New species of Myrmosinae (Hymenop- tera: Mutillidae) from Southeastern Asia. Zootaxa 4656(3): 525-534. Kumar GP, Lelej AS, Das D, Raveendran HKP, Loktionov VM (2019) Discovery of the genus Kudakrumia Krombein, 1979 (Hymenoptera: Mutillidae) in India and description of a new species. Zootaxa 4612(6): 260-266. Loktionov VM, Lelej AS (2019) Contribution to the knowledge of the subgenus Stigmatodipogon \shikawa, 1965 of the genus Dipogon Fox, 1897 (Hymenoptera: Pompilidae). Zootaxa 4613(2): 363-368. Ilyasov RA, Youn HG, Lee M-l, Kim KW, Proshchalykin MYu, Lelej AS, Takahashi J-I, Kwon HW (2019) Phylogenetic relationships of Russian Far-East Apis cerana with other North Asian populations. Journal of Apicultural Science 63(2): 289-314. Loktionov VM, Lelej AS, Liu J-X (2019) A new genus of spider wasps (Hymenoptera, Pompilidae) from China. Far Eastern Entomologist 376: 1-14. Loktionov VM, Lelej AS (2019) The spider wasp genus Teélostholus Haupt, 1929 (Hy- menoptera, Pompilidae) in Indonesia, with the description of three new species. Zootaxa 4648(2): 260-272. 2020 Engel MS, Lelej AS (2020) On the spelling of family-group names based on the genus Trigonalys Westwood (Hymenoptera: Trigonalidae versus Trigonalyidae). Ento- mologist’s Monthly Magazine 156(1): 58-60. Gadallah NS, Lelej AS, Brothers DJ (2020) Biodiversity of the aculeate wasps (Hy- menoptera: Aculeata) of the Arabian Peninsula: Vespoidea, Mutillidae. Zootaxa 47 54( i) 14 1-152. Lelej AS, Terine JB, Kumar GP, Das D, Sureshan PM (2020) Review of the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae) of the Indian subcon- tinent, with description of two new species. Zootaxa 4822(2): 191-208. Lelej AS (2020) To the knowledge of velvet ants of the genera Artiotilla Invrea, Ra- doszkowskitilla Lelej and Taiwanomyrme Tsuneki (Hymenoptera: Mutillidae). Far Eastern Entomologist 410: 1-10. Lelej AS (2020) Three new species of velvet ants from the tribe Smicromyrmini (Hy- menoptera: Mutillidae) with unusual color syndrome. Far Eastern Entomologist 47 J=9. Pagliano G, Brothers DJ, Cambra R, Lelej AS, Lo Cascio P, Matteini Palmerini M, Scaramozzino PL, Williams KA, Romano M (2020) Checklist of names in Mu- tillidae (Hymenoptera), with illustrations of selected species. Bollettino del Museo Regionale di Scienze Naturali 36(1—2): 5-425. Editorial 7s 2021 Okayasu JA, Lelej AS, Williams KA (2021) Review of the genus Fotrogaspidia Lelej, 1996 (Hymenoptera: Mutillidae: Trogaspidiini). Zootaxa 4920(1): 56-90. Sutanto DD, Williams KA, Nugroho H, Lelej AS (2021) To the knowledge of the vel- vet ant Genus Bischoffitilla Lelej (Hymenoptera: Mutillidae) in Indonesia. Treubia 48(1): 1-12. In these works, 3 tribes, 1 subtribe, 1 genus, 41 species and 1 subspecies of Hyme- noptera from 4 families are described in total as new to science. The whole list of the new taxa described by Arkady Lelej therefore includes 364 names. List of the hymenopteran taxa described by Arkady Lelej (2017 — June 2021) Family Mutillidae Apteromutillini Brothers & Lelej, 2017 Ctenotillini Brothers & Lelej, 2017 Dasymutillini Brothers & Lelej, 2017 Euspinoliina Brothers & Lelej, 2017 Eotrogaspidia adhabar Okayasu, Lelej & Williams, 2021 Eotrogaspidia buddha Okayasu, Lelej & Williams, 2021 Eotrogaspidia hauseri Okayasu, Lelej & Williams, 2021 Eotrogaspidia melanopleura Okayasu, Lelej & Williams, 2021 Ephucilla hejunhuai Lelej, 2020 Krombeinella brothersi Williams, Lelej & Thaochan, 2019 Kudakrumia loaibonho Williams, Lelej & Thaochan, 2019 Kudakrumia malaenglek Williams, Lelej & Thaochan, 2019 Kudakrumia rangnekari Girish Kumar & Lelej, 2019 Odontomutilla fletcheri Lelej, Terine & Girish Kumar, 2020 Odontomutilla sairandhriensis Terine, Girish Kumar & Lelej, 2020 Pherotilla brothersi Zhou, Lelej & Xu, 2017 Promecidia abnormis Lelej, 2016 Promecidia chui Lelej et Xu, 2016 Promecilla ovtchinnikovi Lelej, 2020 Sinotilla nigrothoracica Okayasu, Williams & Lelej, 2018 Smicromyrme burgeri Lelej, 2020 Taiwanomyrme cheni Lelej, 2020 Zavatilla nepalensis Zhou et Lelej, 2018 Zavatilla xuzaifui Zhou, Lele} et Williams, 2018 Zeugomutilla pangi Lelej, 2017 Family Pompilidae Eopriocnemis Loktionov & Lelej, 2019 8 M.Yu. Proshchalykin & V.E. Gokhman / Journal of Hymenoptera Research 84: 1-9 (2021) Dipogon lao Loktionov & Lelej, 2018 Dipogon wasbaueri Loktionov & Lelej, 2019 Dipogon siam Loktionov & Lelej, 2019 Eopompilus pseudominor Loktionov, Lelej & Xu, 2017 Eopriocnemis albiflagellata Loktionov & Lelej, 2019 Morochares sinica Loktionov & Lelej, 2018 Morochares wahisi Loktionov & Lelej, 2018 Morochares wolfi Loktionov & Lelej, 2018 Morochares xuzaifui Loktionov, Lelej & Liu, 2018 Nipponodipogon gusenleitnerorum Loktionov & Lelej, 2018 Nipponodipogon orientalis Loktionov, Lelej & Xu, 2017 Nipponodipogon shimizui Loktionov, Lelej & Xu, 2017 Priocnemis yunnanensis Loktionov, Lelej & Xu, 2017 Sphictostethus arfak Loktionov & Lelej, 2018 Sphictostethus papua Loktionov & Lelej, 2018 Stigmaporus volgadon Loktionov & Lelej, 2016 Telostholus celebes Loktionov & Lelej, 2019 Telostholus rinjani Loktionov & Lelej, 2019 Telostholus sulawesi Loktionov & Lelej, 2019 Family Apidae Apis cerana ussuriensis llyasov, Takahashi, Proshchalykin, Lelej & Kwon, 2019 Family Sierolomorphidae Sierolomorpha trjapitzini Mokrousov & Lelej, 2018 In addition to his talent as a scientist, Arkady possesses a number of other remark- able features. First, he is kind and respectful to all surrounding people, including colleagues at work and people on the street. His wonderful politeness and ability to avoid conflicts, i.e., features which are definitely rare and needed nowadays, allow him to quickly establish connections with unfamiliar people who are always positively impressed by their first meeting with him. We know that Arkady’s personality and proficiency in insect collecting, are highly valued by his colleagues (in total, 92 ento- mologists from 20 countries, including authors of this Special Issue), who named 103 (!) insect taxa in Arkady’s honor (COLEOPTERA: Salpingidae (1); DIPTERA: Asi- lidae (5), Cecidomyiidae (1), Chironomidae (1), Dolichopodidae (1), Hybotidae (2), Phoridae (1), Rhagionidae (2), Syrphidae (4); HEMIPTERA: Aphilidae (1); HYME- NOPTERA: Andrenidae (2), Apidae (1), Aulacidae (1), Bethylidae (3), Braconidae (14), Bradynobaenidae (1), Cephidae (1), Chrysididae (3), Colletidae (2), Crabroni- dae (6), Cynipidae (1), Diapriidae (2), Drynidae (2), Encyrtidae (2), Eulophidae (1), Eurytomidae (2), Formicidae (3), Halictidae (1), Ichneumonidae (8), Mutillidae (5), Mymaridae (1), Pamphiliidae (1), Peleserphidae (2), Pompilidae (4), Proctotrupidae (1), Pteromalidae (3), Sphecidae (1), Tenthredinidae (1), Torymidae (1), Vespidae (3); Editorial 9 LEPIDOPTERA: Sesiidae (1), Uraniidae (1); ORTHOPTERA: Acrididae (1), Grylli- dae (2). We believe we can hardly find similar examples among modern entomologists, and we feel privileged to know him as one of the great modern entomologists, a friend, and a delightful human being. The Editors and the Editorial Office of the Journal of Hymenoptera Research, numerous colleagues acting as reviewers of the papers, as well as the Administration of the Federal Scientific Center of the East Asia Terrestrial Biodiversity, were of great help in preparing this Special Issue. Happy 75" birthday, Arkady Stepanovich! We wish you a long and happy life and many new amazing discoveries!